Appellant, Michael Smith, appeals his conviction for carrying a pistol without a license (“CPWL”), possession of an unregistered firearm, unlawful possession of ammunition, and unlawful possession of a firearm.1 He asserts that the trial court committed “reversible error” in excluding his expert testimony on relevancy grounds, and that the court violated his Sixth Amendment right to present a defense. For the reasons stated below, we are constrained to reverse the trial court’s judgment and remand this case for a new trial.
FACTUAL SUMMARY
The government presented evidence showing that on May 2, 2008, Metropolitan Police Department (“MPD”) Officers Dioni Fernandez, Thomas Sheehan, and Matthew Jones responded to a “radio run” for a six foot tall black male with a gun, wearing jeans and a white t-shirt and located in an apartment complex in the 6600 block of Georgia Avenue, in the Northwest quadrant of the District. Officer Fernandez entered the apartment complex first, •with Officers Sheehan and Jones following immediately behind him. Upon entry, Officer Fernandez saw Mr. Smith amidst a group of five or six people. He was holding, perhaps sitting on or straddling, a bicycle. Mr. Smith was the only person in the group who matched the description given on the radio run, and upon seeing the officers, he dropped the bike and began walking away “at a fast pace” while holding his waistband. He ignored Officer Fernandez’s initial “commands to show his hands.”
Officer Sheehan saw Mr. Smith “walking briskly ..., with his hand in his waistband[,] ... looking over his shoulder.” He ordered him to stop approximately five times, and then he stopped Mr. Smith and searched him, but the search produced no weapons. Officer Fernandez returned to the bike that he saw Mr. Smith holding and found “a pouch underneath the seat.” He touched the pouch, felt what he believed was a gun, unzipped the pouch, and saw a gun handle. Officer Fernandez informed Officer Sheehan about the gun. Officer Sheehan transported Mr. Smith “off the scene.” Officer Fernandez rode the bike, with the gun zipped in the pouch, to the nearby Fourth District police station for processing.
Officer Holly Paige of MPD’s Crime Scene section, Forensic Science Division, used an ultraviolet blue powder2 to process the gun for fingerprints on the night of May 2, 2008. The test revealed one fingerprint. Officer Paige was unable to process the bike for fingerprints because Officer Fernandez contaminated it when he rode it to the police station. Officer Paige swabbed the gun for DNA, but never sent the swab for processing because the procedure was costly. MPD fingerprint specialist Barbara Evans also testified as an expert -witness in “the examination of latent prints.” In her thirty-six years with the FBI and MPD, she examined “at least 10,000 or more” latent fingerprints. Ms. Evans examined the fingerprint that Officer Paige recovered and stated that it was not usable because it did not contain enough identifiable points or *1192characteristics to be successfully matched to, or exclude, Mr. Smith. Officer Paige testified that in her experience only five to ten percent of fingerprints recovered from guns are usable.
During cross-examination, Officer Paige admitted that the District of Columbia’s MPD had issued a Special Order3 requiring officers to use the Superglue4 fuming method to recover latent fingerprints from guns that were not found on the defendant’s person.5 She was asked whether the Special Order stated that: “Where a firearm is not found on the defendant’s person, the CSSO ... shall ... us[e] the [S]uperglue method (unless the Forensic Sciences Lab personnel determine that another method would have a better likelihood of success because of the surface involved).”6 Officer Paige responded: “Correct.” She acknowledged that the MPD had Superglue available as well as “mobile units that can come to the case to perform Super[g]lue fuming.” On re-direct, Officer Paige stated that she preferred to use the powder method.
Mr. Smith intended to call his own fingerprint expert, Mr. Wynn, to rebut Officer Paige’s and Ms. Evans’s testimony. On August 19, 2008, the defense provided notice to the government of its intent to call Mr. Wynn “as an expert in the lifting, analysis, and comparison of fingerprints.” The government made an oral motion to exclude Mr. Wynn’s testimony. Mr. Smith’s written Opposition to the government’s motion indicated, in part, that:
Mr. Wynn would testify that [S]uperglue fuming is a superior method to the powder method used in this case by the Crime Scene Search Officer (CSSO). The fact that the police did not use a method available to them is an argument that the jury should acquit based on the government’s lack of evidence. Further, to the extent the government presented the CSSO as a trained professional who did her [investigation] adequately and sufficiently, the defense is entitled to be able to put Mr. Wynn on to rebut that inference that the CSSO is a witness who the factfinder can credit and believe that she did her job adequately. Mr. Wynn would allow Mr. Smith to argue by way of substantive, affirmative evidence that the CSSO’s work is a reason to doubt the government’s case.
In addition, the Opposition stated:
Mr. Wynn would testify that the choice of the powder method utilized in this *1193case decreased the likelihood of recovering a latent print that would be of value to either build a case of conviction against Mr. Smith or to exclude him as having left a print. Mr. Wynn would also testify about recovery rates of latent prints on different porous surfaces, particularly guns. Mr. Wynn would also testify as to the likelihood of recovering a latent print from a bicycle using the [S]uperglue fuming method and other techniques.
Mr. Smith’s Opposition further emphasized that “[t]estimony about the methods used by the [MPD] and the methods they had available [would] go[ ] to the quality of the police investigation!,]” and would be “relevant to the contested issue of whether Mr. Smith possessed a gun.” [Id.] He also stressed that Mr. Wynn would address areas about which Ms. Paige and Ms. Evans testified:
Further, and importantly, these areas: recovery rates, comparison of latent fingerprints for both inclusion and exclusion purposes, and recovery of fingerprints from certain surfaces, the dusting of a bike for fingerprints were all areas of testimony by the [g]overnment’s witnesses .... Mr. Smith now simply seeks to put on his own expert in his defense in these and related areas.
The government objected to Mr. Wynn’s testimony “on relevancy grounds” and stated that it would be cumulative. The prosecutor argued that Mr. Smith was able to elicit all of Mr. Wynn’s proffered testimony during his cross-examination of Officer Paige and Ms. Evans.
The parties and the trial court discussed the issue of Mr. Wynn’s testimony. The trial judge indicated that he needed to do “additional research” before deciding whether to allow Mr. Wynn’s testimony. The following day, the trial court relied on Jackson v. United States, 768 A.2d 580 (D.C.2001), to exclude Mr. Wynn’s testimony as irrelevant and unhelpful in aiding the jury “in determining any issue in this case.” The trial court concluded that (1) Mr. Wynn’s testimony was irrelevant because the fact that the fingerprint was useless was undisputed; (2) Mr. Wynn’s testimony would not aid the jury in determining any of the issues in dispute; (3) Mr. Smith was able to establish his theory of defense sufficiently through cross-examination, and thus; (4) there was nothing Mr. Wynn’s testimony would add to Mr. Smith’s case or to “the defense theory of the ease” or to any matter that was “outside of a juror’s ability to understand and hear this case.” In sum, the trial court reiterated that: “there [was] nothing [Mr. Wynn] could add, with respect to what has been placed on the record, and with respect to the defense theory of the case, that would assist the jury.”
Defense counsel took issue with the trial court, stating in part:
Mr. W[y]nn would add substantial testimony, with respect to the very things that the Government’s witnesses testified about. Ms. Evans was asked about exclusion, dissimilarity, if there’s a usable print.
Mr. W[y]nn would be able to amplify that answer, and we want to present affirmative evidence, the Sixth Amendment, with respect to one point of dissimilarity being able to exclude the person.
She talked about patterns, and didn’t give a straight — didn’t give an answer that Mr. W[y]nn will give, with respect to that point. That makes Mr. W[y]nn’s testimony relevant to Ms. Evans’s testimony.
What is relevant, Your Honor, for reasonable doubt, the defense theory has *1194been lack of investigative efforts, lack of work. I’m not maintaining that the print that was going to be found would exclude Mr. Smith or be dissimilar. It may even convict him. And Mr. W[y]nn would testify that if a usable print were found, it may be matched to him.
And, here, in this case, the jury is going to have no idea what Super[g]lue fuming is.
He will testify that had proper methods been used, [the print] may have had value.
Well, the — the issue of the case is the investigative work by the police. His testimony will help inform the jury, with respect to both the investigation and quality and quantity of the work they did, and the lack of quality and quantity. Mr. W[y]nn is familiar with MPD protocols, familiar with their resources, and he can speak to what they could have done. He’s walking reasonable doubt, if I’m allowed to put him there.
The trial court was unpersuaded and said to defense counsel:
[J]ust as the [c]ourt pointed out in the Jackson case, you’ve put that issue before the jury by cross-examining the Crime Scene Search officer, by cross-examining Officer Fernandez about the responsibilities that are set forth in the Metropolitan Police Department general orders, with respect to maintaining and protecting the integrity of evidence. You’ve put that before the jury. You could make that argument. You’ve got facts upon — in the record, from which you can legitimately make that argument. I don’t see that Mr. W[y]nn’s testimony is going to aid the trier of fact, in any way. And, indeed, actually, I think it’s going to confuse the jury, more than anything else.
ANALYSIS
Mr. Smith argues that, by excluding Mr. Wynn’s testimony as irrelevant, the trial court abused its discretion and violated his Sixth Amendment right to present a defense. He contends that “[t]he trial court’s misapplication of Jackson and subsequent exclusion of [Mr.] Wynn[’s testimony] violated [his] constitutional rights to due process and to an opportunity to present a complete defense, as well as the rules of evidence.” He claims that Jackson supports his argument that Mr. Wynn’s testimony was relevant.7 He states that the government impermissibly “benefitted from the testimony,” particularly Ms. Evans’s statement that the fingerprint was “insufficient for all comparison purposes, including for the purpose of attempting to exclude [Mr. Smith] from the print,” and Officer Paige’s testimony that only five to ten percent of fingerprints retrieved from guns are usable. He maintains that the probative value of Mr. Wynn’s testimony outweighed any prejudice that the government might assert.
Furthermore, Mr. Smith asserts that the trial court excluded evidence relevant to his defense. He points to the government’s closing argument, during which the prosecutor stated that the sole reason for calling Officer Paige and Ms. Evans as witnesses was to prove that a lack of fingerprint evidence was an unreasonable basis for finding Mr. Smith not guilty. He *1195maintains that he had no opportunity to refute the government’s statement with testimony from his own expert.8 Mr. Smith also argues that because the government’s case against him was weak, Officer Paige’s and Ms. Evans’s testimony “was critical to bolstering the government’s thin case against [him],” and that “[w]ithout defense testimony challenging the police officers’ assertions, the government unfairly benefitted from the unimpeached testimony that the police officers did all that they could to find the true owner of the gun.”
The government argues that the trial court did not abuse its discretion in excluding Mr. Wynn’s testimony and that, even assuming error, it was harmless. It maintains that Mr. Wynn’s testimony would not have aided the jurors, only confused them, and that Mr. Smith had ample opportunity to present his defense through cross-examination. It contends that “there was no evidence to suggest that had the Super[g]lue method been used, a print of value would have been recovered.” And “even if Mr. Wynn could have demonstrated that the Super[g]lue method generally produced a higher rate of recovery ... to suggest that the result would have been any different in this case [had the Superglue method been used] would have encouraged the jury to speculate on theoretical possibilities rather than decide the case based on the evidence actually before it.” Finally, the government argues that even if Mr. Wynn’s testimony was relevant to Mr. Smith’s “lack of investigation” theory, it was nevertheless inadmissible because its probative value was outweighed by its potential prejudicial effect.9
The decision to admit or exclude expert testimony is within the trial court’s sound discretion. Dyas v. United States, 376 A.2d 827, 831 (D.C.1977). However, “‘the discretion generally vested in the trial court with respect to the admissibility of evidence must be weighed against the constitutional rights of the defendant to confront witnesses and to present a defense.’ ” Clark v. United States, 639 A.2d 76, 81 (D.C.1993) (quoting Bassil v. United States, 517 A.2d 714, 716 (D.C.1986) (citation omitted)); accord, Russell v. United States, 17 A.3d 581, 585 (D.C.2011) (citing Howard v. United States, 656 A.2d 1106, 1117 (D.C.1995) (other citations omitted)). Generally, expert testimony should be admitted if it is relevant and “likely to aid the trier of fact in the search for the truth.” Dyas, 376 A.2d at 831. “Evidence is relevant if it makes the determination of [an] action more or less probable than it would be without that evidence.” Dockery v. United States, 746 A.2d 303, 306 (D.C.2000). “[T]he decision to exclude expert evidence must be made on a case-by-case basis, grounded on the proffer made and on its potential to assist the jury in the particular case before the court.” Benn v. United States, 978 A.2d 1257, 1273 (D.C.2009) (citing Green v. United States, 718 A.2d 1042, 1051 (D.C.1998)).
We have long followed the test articulated in Dyas, supra, for the admission or exclusion of expert testimony:
*1196(1) the subject matter must be so distinctively related to some science, profession, business or occupation as to be beyond the ken of the average layman;
(2) the witness must have sufficient skill, knowledge or experience in that field or calling as to make it appear that his opinion or inference will probably aid the trier in his search for truth; and (3) expert testimony is inadmissible if the state of the pertinent art or scientific knowledge does not permit a reasonable opinion to be asserted even by an expert.
Id. at 832 (quoting McCormiok on EvidenCE, § 13 at 29-31 (E.Cleary, 2d ed.1972)) (internal quotation marks omitted) (alteration in original). Moreover, consistent with the Dyas test, we have held that expert testimony concerning the feasibility of recovering fingerprint evidence can be relevant to a defendant’s defense theory. Jackson, supra, 768 A.2d at 590 (exclusion of testimony was error “because Jackson’s defense was that his wrongful arrest stemmed from the officers’ neglect ... of their duty to gather available fingerprint evidence”). “[T]he relevance of [fingerprint expert] testimony to that defense [i]s heightened” once the government has presented evidence on the futility of gathering fingerprint evidence. Jackson, 768 A.2d at 590.
Whether trial court evidentiary error in admitting or denying expert opinion testimony is harmless is governed in this case by the non-constitutional standard announced in Kotteakos v. United States, 328 U.S. 750, 765, 66 S.Ct. 1239, 90 L.Ed. 1557 (1946). Reversal may or may not be required under the Kotteakos standard:
If, when all is said and done, the conviction is sure that the error did not influence the jury, or had but very slight effect, the verdict and the judgment should stand, except perhaps where the departure is from a constitutional norm or a specific command of Congress. But if one cannot say, with fair assurance, after pondering all that happened without stripping the erroneous action from the whole, that the judgment was not substantially swayed by the error, it is impossible to conclude that substantial rights were not affected....
Id. at 764-65, 66 S.Ct. 1239 (citation and footnote omitted); see also Russell, supra, 17 A.3d at 589 [ (citing Kotteakos, supra, 328 U.S. at 765, 66 S.Ct. 1239) ] (we cannot say “ ‘with fair assurance ... that the judgment was not substantially swayed by the error’ ”).
We believe that Mr. Smith has the better of the arguments in this particular case. Here, Mr. Smith sought to present testimony from Mr. Wynn, a fingerprint expert with thirty-five years experience who had testified “numerous times” in the trial court and who was familiar with “MPD methods, resources, and protocols.” Counsel for Mr. Smith explained in writing and orally during trial on February 5, 2009, that Mr. Wynn’s testimony was relevant because it addressed whether “the government has met its burden of proof beyond a reasonable doubt”; and that he would provide testimony concerning the quality and adequacy of the government’s investigation and the superiority of the available Superglue fuming method in lifting fingerprints.10
Mr. Wynn’s proffered testimony concerned the success rate of lifting usable prints off of guns when utilizing the Superglue method. According to Mr. Wynn’s *1197proffered testimony, closer to fifty percent of usable fingerprints are recovered when utilizing Superglue fuming, which contradicted Officer Paige’s five to ten percent statistic. In addition, Mr. Wynn would have testified that (1) the Superglue method could have yielded fingerprints off of the contaminated bike, which Officer Paige stated was not possible; and (2) a defendant may be excluded with less than the eight identifiable characteristics that Ms. Evans testified were necessary for exclusion. As such, Mr. Wynn’s testimony was directly relevant to the defense theory that the police investigation was deficient in that Ms. Paige did not use the superior Superglue method that was available.11 Furthermore, Mr. Wynn’s testimony was relevant to assist the jury in determining whether a reasonable doubt existed as to whether Mr. Smith possessed the gun found in the pouch under the seat of the bike. Moreover, the government expressly relied on Officer Paige’s and Ms. Evans’s testimony to rebut Mr. Smith’s defense. It explicitly argued to the jury that the low success rate of gathering useful prints and uselessness of the recovered print in Mr. Smith’s case should cure any reasonable doubt the lack of physical evidence may have created in the jurors’ minds. But the jurors, whose function it “is to independently consider the evidence and attach a relative weight to each item of evidence,” Wheeler v. United States, 930 A.2d 232, 243 (D.C.2007), did not have the benefit of testimonial evidence from Mr. Smith’s fingerprint expert. In declaring that Mr. Wynn’s testimony was irrelevant because the fingerprint lifted by Ms. Paige was useless, the trial court apparently inadvertently missed the essence of defense counsel’s argument about the superiority of the Superglue method in lifting latent fingerprints, the inadequacy of MPD’s investigation, and the relationship of the lack of that evidence to a juror’s reasonable doubt.
Furthermore, a review of the record calls into question the trial court’s exclusion of Mr. Wynn’s testimony on the ground that Mr. Smith was able to establish his theory of defense through cross-examination and on the ground that there was nothing in Mr. Wynn’s testimony that would add to his case or aid the jury in determining the issues in dispute. The record reveals that Mr. Wynn’s proffered testimony was not elicited during Officer Paige’s or Ms. Evan’s cross-examination. Nor did Officer Paige ever fully explain the Superglue method or its significance. Without an explanation of the Superglue method and its rate of success in retrieving fingerprints, as Mr. Wynn was prepared to explain, the significance of Officer Paige’s decision to ignore the MPD directive and use the powder method was not brought to the jury’s attention. Not only could Mr. Wynn’s testimony have undermined the government’s argument, it likely would have impacted Officer Paige’s and Ms. Evan’s credibility as witnesses. And, rather than confuse the jury, Mr. Wynn’s testimony would have explained to the jury that there were other methods, superior methods, that MPD could have used to lift latent fingerprints from the gun and the bike in an effort to determine whether to exclude or include Mr. Smith with respect to the gun.
In addition, the government’s case against Mr. Smith was not particularly strong or overwhelming. He was charged with CPWL, unlawful possession of ammunition, and unlawful possession of a fire*1198arm. The entirety of the government’s evidence linking Mr. Smith to the gun recovered from the bike was Officer Fernandez’s testimony that he saw Mr. Smith holding the bike on which he found the gun in a pouch underneath the seat, and that testimony was partially impeached.12 The other responding officers witnessed Mr. Smith in a group of five or six persons, with at least one other person holding a bike. Neither of those officers saw him holding or even standing near the bike on which the gun was found; they only saw him walking away from the area where the bike was located. Even the trial court noted the weakness of the government’s case.13
In light of these circumstances, we are constrained to hold that the trial court abused its discretion by excluding the testimony of Mr. Wynn. See Johnson v. United States, 398 A.2d 354 (D.C.1979) (discussing abuse of discretion). Under Dyas, Mr. Wynn’s proffered expert testimony was related to the science of lifting fingerprints from objects, particularly the Superglue method specified in the MPD order, as well as the adequacy of the government’s investigation; and Mr. Wynn’s proffered testimony also concerned the points and characteristics of latent fingerprints, subjects addressed by government witnesses Paige and Evans. Hence, the proffered testimony not only was clearly relevant, but it also was beyond the ken of the average layperson. In addition, there was no question about Mr. Wynn’s skill, knowledge, and experience given his thirty-five years as a fingerprint expert and his prior testimony in the Superior Court. Furthermore, as we said in Jackson, supra:
[The defendant’s] defense was that his wrongful arrest stemmed from the [police] officers’ neglect, at least of their duty to gather available fingerprint evidence. The relevance of [the defense expert’s] testimony to that defense was heightened once [the government’s expert] was allowed to testify that, in the view of police experts, fingerprint analysis was futile in cases such as this. Viewed purely as a question of admissibility, therefore, we think the court erred in denying [the defendant’s] request to call [a fingerprint expert] to testify.
Id. at 590.
The remaining question is whether the trial court’s exclusion of Mr. Wynn’s testimony was harmless. We are convinced that the exclusion was not harmless under the non-constitutional Kotteakos standard.14 The government’s case against Mr. Smith was not strong or overwhelming, as we have indicated and as the trial court recognized. In addition, Officer Paige’s and Ms. Evans’s limited expertise with respect to alternative methods of lifting fingerprints prevented Mr. Smith from presenting his evidence through their *1199cross-examination. Expert testimony concerning the inadequacy of the government’s fingerprint investigation and the availability of another, superior method for lifting fingerprints might well have created a reasonable doubt in the minds of jurors. See Greer v. United States, 697 A.2d 1207, 1210 (D.C.1997) (“Reasonable doubt is a doubt arising from the evidence, or from lack of evidence, after consideration of all the evidence”). Consequently, we “cannot say with fair assurance ... that the [trial court’s] judgment was not substantially swayed by the error”; indeed, “[w]e think it highly probable that the error had substantial and injurious effect or influence in determining the jury’s verdict.” Kotteakos, supra, 328 U.S. at 765, 776, 66 S.Ct. 1239; Jackson, supra, 768 A.2d at 590.
Accordingly, we are constrained to reverse Mr. Smith’s conviction and remand this case for a new trial.
So ordered.